Emily "Eve" Best (born 31st July 1971) is an English actress, best known for her roles as Dr. O'Hara in the Showtime television series Nurse Jackie, also as Wallis Simpson in the 2010 film The King's Speech, and Dolley Madison in the 2011 American Experience television special about that First Lady. Eve Best grew up in Ladbroke Grove, London and attended Wycombe Abbey Girls’ School before going on to Lincoln College, Oxford, where she read English. Among her earliest public performances were with the W11 Opera children's opera company in London at the age of nine. After graduating from Oxford where she had appeared in OUDS productions, and toured to the Edinburgh Festival, she made her professional debut as Beatrice in Much Ado About Nothing at the Southwark Playhouse.
After a period working on the London fringe, Best trained at the Royal Academy of Dramatic Art (RADA) in London. After graduating in 1999 she appeared in a revival of 'Tis Pity She's a Whore at the Young Vic for which she won both the Evening Standard and Critics' Circle best newcomer awards; she adopted her grandmother's name as a stage name, as an Emily Best was already registered with British Actors' Equity Association.
Best won an Olivier Award for playing the title role in Hedda Gabler and was nominated for the same award the following year for her performance as Josie in Eugene O'Neill's play A Moon For The Misbegotten at the Old Vic Theatre in London.
In early 2007, she starred in a Sheffield Crucible production of As You Like It which played for a short time at the RSC's Swan Theatre in Stratford as part of their Complete Works season. In the same year she performed in the Broadway transfer of A Moon For The Misbegotten for which she was nominated for a Tony Award as Best Actress in a Play.
Best appeared in Harold Pinter's The Homecoming at the Cort Theatre in New York, which co-starred Ian McShane, Raúl Esparza and Michael McKean. Daniel Sullivan directed the 20-week limited engagement, which ran until 13 April 2008.
Television appearances include Prime Suspect: The Final Act (2006), Waking the Dead (2004), Shackleton (2002), and The Inspector Lynley Mysteries (2005).
She appears as Lucrece in the Naxos audiobook version of Shakespeare's The Rape of Lucrece. She also starred in a 2000 BBC Radio 4 production of Emma.